Description
Key Characters:
Growth Form:
Dioecious shrubs 2–6 dm tall.
Stems:
Stems often swollen at the nodes, terete (cylindrical), 4–angled or –ridged, numerous from the base ascending to erect, glabrous, internodes 0.5–2.3 long, deep yellowish green.
Roots:
Fibrous root system.
Leaves:
Leaves simple.
Opposite.
Blades narrowly linear to linear–subulate, weakly falcate, 4–6 (–8) cm long, 0.18–0.25 cm wide.
Apex attenuate.
Base leaf bases often connate around the stem, base slightly tapering.
Surfaces glabrous, slightly fleshy, arched to recurved, bright yellowish green.
Margins thickened and weakly revolute.
Only the midvein evident, the midvein impressed on upper surface.
Sessile.
Stipules absent.
Flowers:
Flowers in inflorescence terminal, with 8–60 flowers, 2.5–5 cm long, laterally contracted, flowers congested, glabrous or densely puberulent with crinkly hairs to 0.25 mm long; bracts green, linear to subulate, those of central axis 5–20 mm long, those of branches and flowers 1.5–2 mm long; pedicels weakly quadrangular, slightly compressed.
Flowers unisexual and plants dioecious.
Calyx of staminate flowers with sepals 2.5–3 mm long, ovate to broadly navicular, subequal, convex at the base, oriented at 80°to 100° angle to the pedicel, green, yellowish green toward the base, often purple toward the apex, opaque, glabrous or puberulent toward the base and margin, margin scarious and usually with a few lacerations toward the apex, apex obtuse. Calyx of pistillate flowers: sepals 2.3–2.8 mm long, ovate to broadly navicular, subequal, the apex often inrolled, convex at the base, oriented at 90° to 110° angle to the pedicel, green, yellowish green toward the base, often purple toward apex, opaque, glabrous or puberulent toward the base and margin, margin scarious and usually with a few lacerations toward the apex, apex obtuse.
Corolla (petals) absent.
Staminate flowers: Stamens 10, the filaments dimorphic, white, the antisepalous whorl 4.2–4.8 mm long, the alternate whorl 3.9–4.5 mm long; anthers 0.5–0.65 mm long, yellow. Pistillate flowers: Stamens (9–) 10, the filaments weakly dimorphic, the antisepalous whorl 0.5–0.8 mm long, the alternate whorl 0.4–0.5 mm long; anthers 0.15–0.35 mm long, yellow, not producing pollen.
Ovary superior (pistillate flowers), 1-celled; ovule placentation free-central; styles 3 (–4). Staminate flowers: styles 3 (–4), short and non–functional.
Fruit:
Capsules 3–4 mm long; narrowly ovoid.
Seeds 0.7–0.8 mm long; orbicular–reniform; compressed; the surface transversely rugose; perisperm starchy; hard or rarely soft; true endosperm absent.
Ploidy:
Habitat:
Extremely rare; known only from arid alpine cliffs.
Elevation Range:
1770–2440 m.